Targeting endothelial progenitor cells (EPCs) for clinical therapeutics, including vascularization in wound healing, has been hindered by the lack of a uniform EPC definition. Various putative EPC populations have been characterized using several genes/surface markers; we hypothesized that overlapping these markers would allow a consensus definition to be validated functionally. Here we examined the overlap of several markers from across the field with EVPs (VE-Cadherin(VECad)+Lineage(Lin)negCD34+CD31lo) in homeostatic murine aorta and an injury model of full-skin excisional wounds, hoping to narrow a consensus definition. Flow cytometry revealed that among markers tested, Protein C Receptor (Procr) and Platelet-derived growth factor receptor alpha (PDGFRα) were consistently more likely to be co-expressed with EVPs (78.04%, 82.28%) than with mature differentiated endothelial (D) cells (VECad+LinnegCD34+CD31+; 54.22%, 24.65%) in the aorta of adult C57Bl/6 mice (n=5; p<0.001). Single-cell RNA-sequencing confirmed clustering of Procr and PDGFRα with EVPs, while other markers studied clustered with D cells. Immunofluorescent staining confirmed Procr+ EVPs co-localized with YFP+ endothelium in the aortae of Cdh5-CreERT2/Rosa-EYFP reporter mice. Aortic Procr+ EVPs had greater endothelial colony forming capability when cultured in vitro (2.74%) than either Procrneg EVPs (0.69%) or Procr+/neg D cells (0%; n=3), and greater engraftment when implanted in vivo (5.67% vs. 0.79%; n=3; p<0.05). Lineage tracing via flow cytometry of developing PDFGRα-MerCreMer/Rosa-EYFP mice showed that aortic YFP+ EVPs at day 0 (D0) differentiated into D cells by D84 (D0 0.74% D cells, D84 4.67%; n=5; p<0.001). This result was then validated in adult full-skin excisional wounds from the same mice, showing that YFP+ EVPs at D0 differentiated into D cells by D5 (D0 0.04%, D5 1.19%; n=7; p<0.005), suggesting that mesenchymal marker PDGFRα marks an EPC population capable of endothelial fate in both homeostasis and injury. The characteristics displayed by Procr+ and PDGFRα+ EVPs suggest these may mark a true EPC population.